I always thought they were put in series to have better control over the tumble rate.
For the left example... If either the carbon or GFO chamber is plugged faster, the flow will increase in the other chamber, causing an undesirable tumble rate.
For the right example... The whole system slows down and it's just time to change the media.
That being said, I'm not sure what pressure the TLF reactors can handle, so that might be a consideration as well.
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