you made no mention of ammonia or nitrite levels the tank was skimmerless so the DOCS may have gone toxic i would do a 3 gallon water change every day for a month to see if things improve if you do the math that works out to a 100% water change in a month i currently change out 28 gallons a week in my tank and that works out to 60% a month i do 2, 14 gallon water changes a week let us know how it pans out
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always just 2cents until im broke
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